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A V-ATPase proton pump rises in stacked tiers, with a large lobed head on top joined by a narrow central stalk to a rotary membrane ring below. The ring is built from twisted rope-like helices wrapped around the base, while a long peripheral arm runs down one side and knobby clusters bulge from the edges. The tiered, machine-like layout shows how this molecular motor turns. Use it in a cell biology or biochemistry class on proton gradients, or display it with other membrane protein models.

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